Abstract
Peirce’s pragmatic stress on meaning in terms of habits of response is, of course, well known. However, the language in which it is usually expressed tends too often to conflate its epistemic and ontological dimensions, thereby hiding from view its full systematic significance. The following discussion will focus on the emergence of such meanings as epistemic relational structures which embody the characteristics of the dynamics of organism-environment interaction in their very internal structure and which lead outward toward the universe, providing an experienced content which is at once epistemic and ontological.